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Willmott, Keith Richard, Jason Piers Wilton Hall, Griff Kane and Karen Johson
1993. Oxford & Cambridge butterfly expedition to Ecuador 1993. Cambridge Expeditions Journal 28: 13-19, 3 figs. [general]
Willmott, Keith Richard, Jason Piers Wilton Hall and Gerardo Lamas
2001. Systematics of Hypanartia (Lepidoptera: Nymphalidae: Nymphalinae), with a test for geographical speciation mechanisms in the Andes. Systematic Entomology 26(4): 369-399, 24 figs., 1 tab. (October) [new taxa: H. dione disjuncta, H. celestia, H. fassli, H. trimaculata, H. t. autumna, H. cinderella; biogeography; Neotropics]
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