Acknowledgements
The authors are grateful to the two anonymous reviewers for their sound criticism and constructive comments
on the earlier versions of this paper. This study is supported in part by the Japanese Ministry of Education,
Culture, Sports, Science and Technology (MEXT) through the Arctic Challenge for Sustainability II (ArCS-
II; JPMXD1420318865), by the Japan Science and Technology Agency through COI-NEXT JPMJPF2013, by
the Japanese Ministry of Environment through Environment Research and Technology Development Fund
JMEERF20192004, and by the Japan Society for the Promotion of Science (JSPS) through Grants-in-Aid for
Scientific Research JP18H01278, JP19H05702 (on Innovative Areas 6102) and 20H01970. Y.K. acknowledges
support from the JSPS Invitational Fellowship for Research in Japan that supported a sabbatical at the University
of Tokyo and ignited this collaboration, for support from the Research Center for Advanced Technology and
Science at the University of Tokyo and the Israeli Science Foundation (grant 996/20).
Author contributions
S. O. designed the research at the suggestion of H.N. and performed the analyses. S. O. and H.N. wrote the
manuscript with discussion and feedback from Y. K. All authors reviewed the manuscript.
Competing interests
The authors declare no competing interests.
Additional information
Supplementary Information The online version contains supplementary material available at
https:// doi. org/
10. 1038/ s41598- 021- 92548-7
.
Correspondence and requests for materials should be addressed to S.O.
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