Pollen Fossil
|
Reference
|
Age
|
Fossil location
|
Calibration node
|
Prior distribution
|
Myrtales crown age estimate
|
Wang et al., 2009
|
91 Ma
|
n/a
|
Root node calibration (A)
|
Normal
Mean - 91
St. Dev. - 2.5
|
Myrtaceidites mesonesus Cookson & Pike
|
Couper, 1960
|
Paleocene
|
New Zealand
|
All Myrtoideae except Xanthostemoneae and Lophostemoneae (B)
|
Exponential
Mean - 1.03
Offset - 61.7
|
Myrtaceidites tenuis Harris
|
Thornhill and Macphail in review
|
Early Eocene
|
Bass Strait, Tasmania
|
Angophora + Corymbia crown (C)
|
Exponential
Mean - 2.17
Offset - 45.0
|
Myrtaceidites eucalyptoides Cookson & Pike
|
Alley et al., 1996
|
Late Eocene
|
Lake Eyre and Otway Basins, South Australia
|
Eucalyptus crown (D)
|
Exponential
Mean - 0.895
Offset - 33.9
|
Myrtaceidites sp.
|
Thornhill and Macphail in review
|
Early Eocene
|
Pilbara, Western Australia
|
Leptospermeae crown (E)
|
Exponential
Mean - 1.952
Offset - 48.6
|
Myrtaceidites verrucosus
Stover & Partridge
|
Thornhill and Macphail in review
|
Late Eocene
|
Hatfield, New South Wales
|
Australasian Myrteae crown
(F)
|
Exponential
Mean - 0.895
Offset - 33.9
|
Myrtaceidites verrucosus
|
Graham, 1985; Romero and Zamaloa, 1985
|
Middle Eocene
|
Panama and Argentina
|
South American Myrteae crown (G)
|
Exponential
Mean - 3.09
Offset - 37.2
|
Myrtaceidites verrucosus
|
Pocknall, 1982
|
Late Oligocene
|
Pomahaka, New Zealand
|
New Zealand Myrteae crown
(H)
|
Exponential
Mean - 1.47
Offset - 23.03
|
Myrtaceidites eugeniioides Cookson & Pike
|
Harris, 1965
|
Early Eocene
|
Princetown, Victoria
|
Syzygieae + Backhousieae crown (I)
|
Exponential
Mean - 1.952
Offset - 48.6
|
Macrofossil
|
Reference
|
Age
|
Fossil location
|
Calibration node
|
Prior distribution
|
Eucalyptus fruits
|
Hermsen et al., 2010
|
Early Eocene
|
Patagonia, South America
|
Crown eucalypts (J)
|
Exponential
Mean - 1.844
Offset - 51.2
|
Corymbia fruit
|
Rozefelds, 1996
|
Early to Middle Eocene
|
Queensland
|
Crown Corymbia/Angophora (K)
|
Exponential
Mean - 4.744
Offset - 45
|
Paleomyrtinaea princetonensis
Pigg, Stockey & Maxwell
|
Pigg et al., 1993
|
Early Eocene
|
United States of America
|
Crown Myrteae (L)
|
Exponential
Mean - 4.066
Offset - 50.0
|
Tristaniandra alleyi
Peter G. Wilson & Basinger
|
Basinger et al., 2007
|
Middle Eocene
|
South Australia
|
Crown Kanieae (M)
|
Exponential
Mean - 3.09
Offset - 37.2
|
Metrosideros leaves and fruits
|
Pole et al., 2008
|
Miocene
|
New Zealand
|
Crown Metrosideros (N)
|
Exponential
Mean - 2.982
Offset - 20.0
|
Taxon
|
Characters
|
1
|
2
|
3
|
4
|
5
|
6
|
7
|
8
|
9
|
10
|
11
|
12
|
13
|
14
|
Acca sellowiana
|
1
|
1&2
|
3
|
0
|
2
|
0&1
|
1
|
3
|
2
|
3
|
0
|
2
|
2
|
3
|
Acmena divaricata
|
0
|
0
|
7
|
3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
2
|
6
|
Acmena graveolens
|
0
|
0
|
7
|
3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
2
|
6
|
Acmena mackinnoniana
|
0
|
0
|
7
|
4
|
0&6
|
0
|
0
|
0
|
0&1
|
0
|
0
|
2
|
2
|
6
|
Acmena smithii
|
0&1
|
0&1
|
2&7
|
4
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
1
|
2
|
Acmenosperma claviflorum
|
1
|
1
|
3
|
0&1
|
3
|
1
|
1
|
0&1
|
0&1&2
|
0
|
0
|
2
|
2
|
2
|
Actinodium cunninghamii
|
0
|
1
|
0
|
2&4
|
0
|
0
|
0
|
0
|
3
|
5
|
0
|
1
|
1
|
0
|
Agonis flexuosa
|
1
|
1&2
|
1&2
|
1
|
3
|
0
|
0
|
0&2
|
0
|
0&1
|
1
|
1
|
1
|
4
|
Amomyrtus meli
|
1
|
1
|
4
|
0
|
2
|
0
|
0
|
0&3
|
2
|
3
|
0
|
4
|
4
|
2
|
Anetholea anisata
|
1
|
1&2
|
2
|
3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
1
|
1
|
4
|
Angophora costata
|
1
|
1
|
2
|
0&1&3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
4
|
4
|
3
|
Angophora hispida
|
1
|
1
|
2
|
0&1
|
0
|
1
|
2
|
0&3
|
0&1
|
0
|
0
|
4
|
4
|
3
|
Archirhodomyrtus beckleri
|
1
|
1
|
4
|
0&3
|
1
|
0
|
0
|
0
|
2
|
3
|
0
|
3
|
3
|
2
|
Arillastrum gummiferum
|
1
|
1
|
2&6
|
3&4
|
0&3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
1
|
2
|
Asteromyrtus symphyocarpa
|
1
|
1
|
1
|
0
|
3
|
1
|
2
|
0&1
|
0&1
|
0
|
1
|
2
|
2
|
4
|
Babingtonia cunninghamii
|
1
|
1
|
5&6
|
0&1
|
0
|
0
|
0
|
0&1
|
1
|
0
|
0
|
1
|
1
|
1
|
Babingtonia densifolia
|
1
|
1
|
1&6
|
1
|
0
|
0
|
0
|
1
|
0
|
1
|
0
|
0
|
0
|
0
|
Babingtonia jucunda
|
1
|
1&2
|
0&1&5
|
0
|
0
|
0
|
0
|
0&1
|
0
|
0
|
0
|
1
|
1
|
0
|
Babingtonia tozerensis
|
1
|
1
|
6
|
4
|
0&3
|
0
|
0
|
0&1
|
0
|
0
|
0
|
1
|
1
|
3
|
Backhousia citriodora
|
1
|
1
|
2&3
|
0&3
|
0&6
|
0&1
|
0&1
|
0&1
|
0
|
0
|
0
|
2
|
2
|
3
|
Backhousia myrtifolia
|
1
|
1
|
2
|
1
|
0
|
0
|
0
|
0&1
|
0
|
0
|
0
|
2
|
2
|
4
|
Baeckea frutescens
|
1
|
1
|
5
|
0&1
|
0
|
0
|
0
|
0&1
|
0
|
0
|
0
|
1
|
1
|
0
|
Baeckea ovalifolia
|
1
|
1
|
0&1
|
1
|
0&3
|
0
|
0
|
1
|
0
|
0
|
0
|
1
|
1
|
2
|
Beaufortia orbifolia
|
1
|
1
|
2&3
|
0
|
3
|
1
|
1&2
|
0&1
|
0
|
0
|
0
|
3
|
3
|
3
|
Blepharocalyx cruckshanksii
|
1
|
1
|
3
|
3
|
2
|
1
|
1
|
0
|
2
|
3
|
0
|
2
|
2
|
3
|
Blepharocalyx salicifolius
|
1
|
1
|
4
|
0&3
|
2
|
0
|
0
|
0
|
2
|
3
|
0
|
1
|
1
|
2
|
Blepharocalyx tweediei
|
1
|
1
|
4
|
0&3
|
2
|
0
|
0
|
0
|
2
|
3
|
0
|
1
|
1
|
2
|
Callistemon polandii
|
1
|
1
|
3
|
0&3
|
0&3
|
1
|
1
|
0&1
|
0
|
0
|
0
|
2
|
2
|
3
|
Calothamnus validus
|
1
|
1
|
2
|
1&2
|
3
|
0
|
0
|
0&2
|
0
|
0
|
0
|
2
|
1
|
4
|
Calyptranthes concinna
|
1
|
1
|
4
|
3
|
2
|
0
|
0
|
0
|
2
|
1
|
0
|
2
|
2
|
2
|
Calyptranthes pallens
|
1
|
1
|
4
|
3
|
2
|
0
|
0
|
0
|
0
|
1
|
0
|
2
|
2
|
2
|
Calytrix tetragona
|
1
|
1
|
1
|
0
|
0
|
0
|
0
|
3
|
0
|
0
|
0
|
2
|
2
|
4
|
Campomanesia guazumifolia
|
1
|
1
|
4
|
3&4
|
2
|
0
|
0
|
0&1
|
2
|
2
|
0
|
3
|
3
|
2
|
Chamelaucium uncinatum
|
1
|
1
|
0
|
3
|
0
|
0
|
0
|
0
|
3
|
5
|
0
|
4
|
3
|
1
|
Choricarpia subargentea
|
1
|
1
|
2
|
1
|
3
|
0
|
0
|
0
|
0&1
|
0&1
|
0
|
2
|
2
|
4
|
Cloezia floribunda
|
1
|
1
|
1&2
|
1
|
0
|
0
|
0
|
0&3
|
0
|
0
|
0
|
1
|
1
|
4
|
Corymbia variegata
|
1
|
1
|
2
|
1
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
4
|
4
|
4
|
Darwinia fascicularis
|
1
|
1
|
0
|
0&1&3
|
0&3
|
0
|
0
|
0
|
3
|
5
|
0
|
3
|
3
|
1
|
Decaspermum humile
|
1
|
1
|
3
|
0&1
|
1&5
|
0
|
0
|
0
|
2
|
1
|
0
|
2
|
1
|
2
|
Eucalyptopsis papuana
|
1
|
1&2
|
3
|
4
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
1
|
1
|
2
|
Eucalyptus curtisii
|
1
|
1
|
2
|
4
|
3
|
0&1
|
0&1&2
|
0
|
0
|
0
|
0
|
2
|
2
|
3
|
Eucalyptus globulus
|
1
|
1
|
3
|
0&3
|
3
|
0
|
0
|
0
|
1&2
|
0&2
|
0
|
3
|
3
|
2
|
Eugenia uniflora
|
1
|
1
|
3&4
|
3&4
|
0&2
|
0
|
0
|
0
|
2
|
1
|
0
|
2
|
3
|
2
|
Euryomyrtus ramosissima
|
1
|
1
|
1
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
1
|
0
|
4
|
Heteropyxis natalensis
|
1
|
1
|
3
|
0
|
0&3
|
0&1
|
0&2
|
2
|
0&1
|
0&1
|
0
|
2
|
2
|
3
|
Hexachlamys edulis
|
1
|
1
|
4
|
3
|
2
|
0
|
0
|
0&1
|
2
|
3
|
0
|
2
|
2
|
2
|
Homalocalyx aurea
|
1
|
1
|
1
|
0&3
|
3
|
0&1
|
0&2
|
0&1
|
2
|
3&4
|
1
|
3
|
3
|
3
|
Homoranthus darwinioides
|
1
|
1
|
0
|
0
|
0
|
0
|
0
|
0
|
3
|
5
|
0
|
3
|
3
|
0
|
Kjellbergiodendron celebicum
|
1
|
1
|
2
|
3
|
3
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
2
|
3
|
Kunzea capitata
|
1
|
1&2
|
1
|
0&3
|
0
|
0
|
0
|
1
|
0
|
0
|
0
|
2
|
2
|
4
|
Legrandia concinna
|
1
|
1
|
4
|
0&3
|
0&2
|
0
|
0
|
0
|
2
|
3
|
0
|
3
|
3
|
2
|
Leptospermum scoparium
|
1
|
1&2
|
1&2
|
3
|
1&3
|
0
|
0
|
0
|
0
|
0
|
0
|
1
|
1
|
4
|
Leptospermum spectabile
|
1
|
1
|
1
|
3
|
1&3
|
0
|
0
|
1
|
0&1
|
0
|
1
|
2
|
2
|
3
|
Leptospermum trinervium
|
1
|
1
|
1
|
3
|
1&3
|
0
|
0
|
0
|
0&1
|
0
|
1
|
2
|
2
|
4
|
Lindsayomyrtus racemoides
|
1
|
1
|
1
|
3
|
0&3
|
0
|
0
|
0
|
2
|
0&3
|
1
|
1
|
1
|
3
|
Lophomyrtus bullata
|
1
|
1&2
|
4
|
3
|
3
|
0
|
0
|
0
|
2
|
1
|
0
|
2
|
2
|
2
|
Lophostemon confertus
|
1
|
1
|
2
|
3
|
3
|
0
|
0
|
0
|
0
|
0
|
0
|
1
|
1
|
3
|
Luma apiculata
|
1
|
1
|
4
|
3&4
|
2
|
0
|
0
|
0
|
2
|
1
|
0
|
1
|
1
|
2
|
Lysicarpus angustifolius
|
1
|
1
|
2
|
3
|
3
|
0
|
0
|
0&1
|
0
|
0
|
0
|
2
|
1
|
4
|
Melaleuca nesophila
|
1
|
1
|
3
|
3
|
3
|
1
|
1
|
0&1
|
1
|
0
|
0
|
1
|
1
|
2
|
Melaleuca viridiflora
|
1
|
1
|
2&3
|
3&4
|
2
|
1
|
1
|
0
|
0&1
|
0
|
0
|
2
|
2
|
2
|
Metrosideros carminea
|
1
|
1
|
2
|
1
|
0
|
1
|
2
|
0
|
0
|
0
|
0
|
1
|
1
|
3
|
Metrosideros diffusa
|
1
|
1
|
3
|
0&3
|
0&3
|
1
|
1
|
0
|
0
|
0
|
0
|
1
|
1
|
2
|
Metrosideros macropus
|
1
|
1
|
2
|
0&3
|
0&3
|
0&1
|
0&1
|
0&1
|
0
|
0&5
|
0
|
2
|
2
|
3
|
Metrosideros nervulosa
|
1
|
1
|
3
|
0
|
0
|
1
|
1
|
0&3
|
0
|
0
|
0
|
3
|
3
|
3
|
Micromyrtus ciliata
|
1
|
1
|
2
|
0&3
|
0
|
0
|
0
|
0&1
|
0
|
0
|
0
|
1
|
1
|
3
|
Myrceugenia leptospermoides
|
1
|
1
|
4
|
0
|
2
|
0
|
0
|
0
|
2
|
3
|
0
|
1
|
2
|
2
|
Myrciaria vexator
|
1
|
1
|
4
|
3&4
|
3
|
0
|
0
|
0
|
2
|
3
|
0
|
2
|
2
|
2
|
Myrteola nummularia
|
1
|
1
|
4
|
0
|
2
|
0
|
0
|
0
|
2
|
3
|
0
|
2
|
2
|
2
|
Myrtus communis
|
1
|
1
|
4
|
3
|
2
|
0
|
0
|
0
|
2
|
3
|
0
|
1
|
1
|
2
|
Neomyrtus pedunculata
|
1
|
1
|
4
|
0&1&3
|
0&3
|
0
|
0
|
0
|
2
|
1
|
0
|
2
|
2
|
2
|
Ochrosperma lineare
|
1
|
1
|
1
|
2
|
3
|
0
|
0
|
0&1
|
1
|
5
|
1
|
1
|
0
|
5
|
Ochrosperma oligomerum
|
1
|
1
|
1
|
2
|
3
|
0
|
0
|
0&1
|
1
|
5
|
1
|
0
|
0
|
4
|
Osbornia octodonta
|
1
|
1
|
1&3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
2
|
4
|
Pericalymma ellipticum
|
1
|
1
|
1
|
0&1
|
1&3
|
0
|
0
|
1
|
0
|
0
|
1
|
3
|
3
|
4
|
Pileanthus filifolius
|
0
|
0
|
7
|
0
|
0
|
0
|
0
|
0&3
|
0
|
0
|
0
|
5
|
3
|
6
|
Pilidiostigma papuanum
|
1
|
1
|
4
|
4
|
2
|
0
|
0
|
0
|
2
|
3
|
0
|
3
|
3
|
2
|
Piliocalyx concinnus
|
1
|
1
|
2
|
3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
2
|
2
|
Pimenta racemosa
|
1
|
1&2
|
3
|
3
|
0
|
0
|
0
|
0
|
0&1
|
0
|
0
|
1
|
2
|
2
|
Psidium cattleianum
|
1
|
1
|
4
|
4
|
0
|
0
|
0
|
0
|
2
|
1
|
0
|
4
|
4
|
2
|
Psiloxylon mauritianum
|
1
|
1&2
|
3
|
3
|
0&3
|
1
|
1
|
0
|
0
|
0
|
0
|
1
|
1
|
2
|
Qualea grandiflora
|
1
|
0&1
|
4
|
0&4
|
3
|
0
|
0
|
0&3
|
1
|
1
|
0
|
4
|
4
|
1
|
Rhodamnia argentea
|
1
|
1
|
4
|
3
|
2
|
0
|
0
|
0
|
2
|
1
|
0
|
1
|
1
|
2
|
Rhodomyrtus macrocarpa
|
1
|
1
|
4
|
3&4
|
2
|
0
|
0
|
0
|
2
|
3
|
0
|
4
|
4
|
2
|
Sphaerantia chartacea
|
1
|
1
|
3
|
3
|
5
|
0
|
0
|
1
|
1
|
0
|
0
|
1
|
1
|
3
|
Stockwellia quadrifida
|
1
|
1
|
2
|
4
|
3
|
0
|
0
|
0&1
|
0&1
|
0
|
0
|
1
|
1
|
2
|
Syncarpia glomulifera
|
1
|
1
|
2
|
1
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
2
|
2
|
4
|
Syzygium aromaticum
|
1
|
1
|
3
|
3
|
0&3
|
1
|
1
|
0
|
1&2
|
0
|
0
|
2
|
2
|
2
|
Syzygium australe
|
1
|
1
|
2
|
1&3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
1
|
1
|
4
|
Syzygium cumini
|
1
|
1
|
3
|
0&3
|
0
|
1
|
2
|
1
|
0&1
|
0
|
0
|
1
|
1
|
3
|
Syzygium maire
|
1
|
1
|
3
|
0&3
|
0
|
1
|
1
|
0
|
1
|
0
|
0
|
1
|
1
|
2
|
Syzygium samarangense
|
1
|
1
|
2
|
4
|
0
|
0
|
0
|
0
|
0&2
|
0
|
0
|
1
|
1
|
3
|
Syzygium wilsonii
|
1
|
1&2
|
3
|
0
|
3
|
1
|
1
|
0&1
|
0
|
0
|
0
|
2
|
1
|
3
|
Tepualia stipularis
|
0
|
0
|
7
|
3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
3
|
1
|
6
|
Thaleropia queenslandica
|
1
|
1
|
0
|
1
|
0
|
0
|
0
|
0
|
3
|
5
|
0
|
0
|
0
|
0
|
Thryptomene saxicola
|
0&1
|
0&1
|
6
|
4
|
3
|
0
|
0
|
0
|
0
|
0
|
0
|
1
|
1
|
2
|
Tristania neriifolia
|
1
|
1
|
1&3
|
0
|
0
|
0
|
0
|
0&1
|
0
|
5
|
0
|
0
|
0
|
0
|
Tristaniopsis laurina
|
1
|
1&2
|
3&6
|
0&3
|
0
|
0
|
0
|
0
|
0
|
0
|
0
|
1
|
1
|
2
|
Ugni molinae
|
0&1
|
0&1
|
4
|
3&4
|
2
|
0
|
0
|
0&2
|
2
|
3
|
0
|
1
|
1
|
1
|
Uromyrtus australis
|
1
|
1
|
4
|
4
|
2
|
0
|
0
|
0&1
|
2
|
3
|
0
|
1
|
1
|
1
|
Verticordia pennigera
|
1
|
1&2
|
0
|
1
|
0
|
0
|
0
|
0
|
3
|
5
|
0
|
3
|
3
|
1
|
Vochysia hondurensis
|
1
|
1
|
4
|
0
|
3
|
0
|
0
|
3
|
3
|
1
|
0
|
?
|
?
|
?
|
Waterhousea floribunda
|
1
|
1
|
2&3
|
0&3&4
|
0&3
|
0&1
|
0&2
|
0
|
0&1
|
0
|
0
|
1
|
1
|
3
|
Waterhousea hedraiophylla
|
1
|
1
|
2
|
0
|
0
|
0
|
0
|
0
|
0&1
|
0
|
0
|
1
|
1
|
3
|
Xanthomyrtus papuana
|
1
|
1&2
|
2
|
0&3
|
0&3
|
0
|
0
|
0
|
0
|
0&1
|
0
|
2
|
1
|
3
|
Xanthostemon chrysanthus
|
1
|
1
|
2
|
3
|
2
|
1
|
1
|
0
|
0
|
0
|
0
|
2
|
2
|
2
|