P. S. Green Royal Botanic Gardens, Kew, Richmond, Surrey, tw9 3AB, England

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Found in the moist, shaded forest, often beside waterfalls.

N.Is.: Now-Now Valley, R.D.Hoogland 11254 (K, NSW); between Palm Glen and Red Rd, M.Lazarides 8090 (CANB, K, NSW); valley S of Mt Bates, P.S.Green 1390 (A, K).

saxifragoides(C.Presl) P.S.Green
Kew Bull. 48: 618(1993)

Trichomanes saxifragoides C.Presl, Hymenophyllaceae 16, 39 (1843); Gonocormus saxifragoides (C.Presl) Bosch, Verh. Kon. Ned. Akad. Wetensch. Afd. Natuurk. 9 (Hymenophyll. Javan.): 9 (1861).T: Philippine Islands, H.Cuming 256; holo: ?PR or ?PRC n.v.; iso: K. So named from a supposed resemblance to some species of Saxifraga.

Illustrations: D.L.Jones & S.C.Clemesha, Austral. Ferns & Fern Allies 2nd edn, 44, fig. 20j (1981), as Gonocormus saxifragoides; D.L.Jones, Encycl. Ferns 66 (1987), as Trichomanes saxifragoides; P.G.Wilson in G.J.Harden, Fl. New South Wales 1: 34 (1990), as Gonocormus saxifragoides.

Epiphytic or lithophytic fern, often forming large patches. Rhizome very slender, creeping, with simple brown hairs. Fronds: stipe wiry, 0.3–2 cm long, hairy; lamina fan-shaped, subreniform, or almost circular, 0.5–2.5 cm long, delicate, dark-green; segments ±oblong, 1–6 mm long, obtuse, emarginate or bifid. Sori immersed in apices of segments; indusia tubular with broadly dilated mouth; receptacle exserted to 0.5 mm when old.

Small Filmy Fern.

Norfolk Is. Localised, but not rare, even though first discovered only in the 1960s. Also known from Africa to Japan, and Malesia to north-eastern Australia, New Caledonia, Fiji and Samoa.

Found in moist, humid valleys in the National Park.

N.Is.: S side of Mt Bates, P.S.Green 1873 (K).


Hymenophyllaceae 17(1843)
from the Greek kephale (a head) and manos (slender or loose), presumably in allusion to the slender receptacles in the type species

Type: C. atrovirens C.Presl

Terrestrial or lithophytic fern. Rhizomes erect or only shortly creeping, relatively thick. Fronds somewhat clustered on rhizome; stipe distinct; lamina usually only 1-pinnate, with false veins absent; cell walls usually thick. Sori usually at tips of ultimate segments; indusium cup-shaped, bell-shaped or tubular, truncate, entire or dilated; receptacle long-exserted.

A genus of c. 60 species, mostly from the Old World; 2 native species on Lord Howe Is. with 1 also present on Norfolk Is.

G.Bentham, Filicales, Trichomanes, Fl. Austral. 7: 700–704 (1878).

Fronds 7–20 cm long, pinnate; segments broad

1. C. atrovirens

Fronds 15–50 cm long, deeply and narrowly tripinnatifid

2. C. bauerianum

Hymenophyllaceae 18, t. 5(1843)

Trichomanes atrovirens (C.Presl) Kuntze, Bot. Zeitung (Berlin) 5: 371 (1847).T: Philippine Islands, H.Cuming 169; holo: ?PR or ?PRC n.v. The epithet comes from the Latin ater (black) and virens (green), in allusion to the dark colour of the fronds.

[Trichomanes javanicum auct. non Blume: W.R.B.Oliver, Trans. & Proc. New Zealand Inst. 49: 117 (1917)]

Illustrations: D.L.Jones & S.C.Clemesha, Austral. Ferns & Fern Allies, 2nd edn, 45, fig. 221 (1981); P.J.Brownsey & J.C.Smith-Dodsworth, New Zealand Ferns & Allied Pl. 69, fig. 71, t. 16A (1989), as Hymenophyllum atrovirens; S.B.Andrews, Ferns Queensland 197, fig. 16.14A (1990).

Terrestrial or lithophytic fern. Rhizome erect, often elevated on stilt-like roots. Fronds: stipe 1–4 cm long, hairy; lamina somewhat harsh-textured, ±erect, 7–20 cm long, pinnate; pinnae 4–6 mm broad, asymmetrical; veins prominent, protruding beyond the margin as tiny mucros. Sori marginal, free; indusia tubular, 1–2 mm long, slightly dilated; receptacle exserted by 1–6 mm.

Lord Howe Is. Rare, no recent records seen. Also known from Australia (northern Qld), Vanuatu, the Solomon Islands, New Guinea and the Philippines.

L.H.Is.: Soldiers or Deep Ck, 1887, T.Whitelegge (NSW).

This species is very closely related to C. javanicum (Blume) C.Presl and C. boryanum (Kunze) Bosch.

bauerianum(Endl.) P.S.Green
Kew Bull. 48: 618(1993)

Trichomanes bauerianum Endl., Prodr. Fl. Norfolk. 17 (1833); Trichomanes meifolium var. bauerianum (Endl.) Hook., Syn. Fil. 1: 137 (1845); Callistopteris bauerianum (Endl.) Copel., Philipp. J. Sci. 67: 65 (1938).T: Norfolk Island, F.L.Bauer; holo: W; iso: K. Named after the Austrian botanical artist, Frederick Lucas Bauer (1760–1826), who collected on Norfolk Is. in 1804–1805.

[Trichomanes polyanthos auct. non Hook.: F.J.H. von Mueller, Fragm. 7: 121 (1870)]

[Trichomanes apiifolium auct. non C.Presl: G.Bentham, Fl. Austral. 7: 703 (1878); J.H.Maiden, Proc. Linn. Soc. New South Wales 28: 729 (1904)]

Illustration: E.B.Copeland, Philipp. J. Sci. 51: t. 42, fig. 2 (1933), as Trichomanes bauerianum.

Terrestrial or lithophytic fern. Rhizome short, erect, supported by coarse roots, with abundant dark brown, filiform scales and hairs 4–7 mm long. Fronds: stipe 5–25 cm tall, filiform, hairy; lamina ±erect, lanceolate in outline, 10–30 cm long, deeply and abundantly tripinnatifid; pinnae often overlapping; segments linear, 0.5–0.75 mm broad, obtuse; veins not protruding. Sori somewhat immersed in tips of lateral pinnae; indusia conoid, 1 mm long, ±flared at apex; receptacle becoming exserted when fully developed, 2 mm long.

Large Filmy Fern.

Occurs on Norfolk Is., Lord Howe Is.

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