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Servers and client applications

The interface of an email client, Thunderbird
Messages are exchanged between hosts using the Simple Mail Transfer Protocol with software programs called mail transfer agents (MTAs); and delivered to a mail store by programs called mail delivery agents (MDAs, also sometimes called local delivery agents, LDAs). Accepting a message obliges an MTA to deliver it,^[65] and when a message cannot be delivered, that MTA must send a bounce message back to the sender, indicating the problem.
Users can retrieve their messages from servers using standard protocols such as POP or IMAP, or, as is more likely in a large corporate environment, with a proprietary protocol specific to Novell Groupwise, Lotus Notes or Microsoft Exchange Servers. Programs used by users for retrieving, reading, and managing email are called mail user agents (MUAs).
When opening an email, it is marked as "read", which typically visibly distinguishes it from "unread" messages on clients' user interfaces. Email clients may allow hiding read emails from the inbox so the user can focus on the unread.^[66]
Mail can be stored on the client, on the server side, or in both places. Standard formats for mailboxes include Maildir and mbox. Several prominent email clients use their own proprietary format and require conversion software to transfer email between them. Server-side storage is often in a proprietary format but since access is through a standard protocol such as IMAP, moving email from one server to another can be done with any MUA supporting the protocol.
Many current email users do not run MTA, MDA or MUA programs themselves, but use a web-based email platform, such as Gmail or Yahoo! Mail, that performs the same tasks.^[67] Such webmail interfaces allow users to access their mail with any standard web browser, from any computer, rather than relying on a local email client.

Filename extensions

Upon reception of email messages, email client applications save messages in operating system files in the file system. Some clients save individual messages as separate files, while others use various database formats, often proprietary, for collective storage. A historical standard of storage is the mbox format. The specific format used is often indicated by special filename extensions:
eml
Used by many email clients including Novell GroupWise, Microsoft Outlook Express, Lotus notes, Windows Mail, Mozilla Thunderbird, and Postbox. The files contain the email contents as plain text in MIME format, containing the email header and body, including attachments in one or more of several formats.